Jeanne La Roche lives alone with her brother in the great northwestern country. Jacques is a ne'er-do-well and has fallen under the suspicion of the mounted police, two of whom are dispatched to arrest him for robbery. The stolen goods are found in his home. Jeanne is too young to be left in their lonely cabin, so she is taken to the post, where the wife of the proprietor welcomes her and gives her a home. Several years later, Donald McLean wins her for his wife. Meantime Jacques escapes from prison, eludes his pursuers and takes refuge in McLean's home.
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Two prospectors, one the father of Sky "Lightning" Bryce and the other the father of Kate Arnold, find a large gold deposit belonging to an Indian tribe.
The story begins as Tom Mills (Tom Mix) rides off to fight in WWI. Leaving his ranch in the care of his sister Ellen (Carmelita Geraghty) and her husband Ed (Carl Miller) Mills returns from the battlefield two years later to find that his brother-in-law has deserted, and the ranch is in a state of ruin and disrepair.
While in Europe, Chaddie Green, a society girl, discovers that she has been left penniless. She returns to the United States and meets Duncan MacKail, who is equally broke though he owns grainland in the West.
After the train station clerk is assaulted and left bound and gagged, then the departing train and its passengers robbed, a posse goes in hot pursuit of the fleeing bandits.
Claire Adams as a girl forced to marry the man she suspects killed her father. When she refuses, she is virtually kept a prisoner along with kid-brother Frankie Lee until a handsome stranger (Jack Conway) rescues them.
Out of the blue, war is declared, making two otherwise friendly neighbouring nations deadly enemies. Amid the inferno, finding themselves on different sides of the conflict, Elsa and Alexis are forced to call off their engagement.
Movie mogul Thomas H. Ince may well have been the director of The Despoiler as indicated by the credits; but since Ince was known far and wide as a glory-hogger, it's also possible that one of his talented lieutenants wielded the megaphone.